The Autobots and Decepticons find themselves involved in humanity's conflict in the Second World War, with the Autobots aiding the Americans, British, & Red Armies battling the Decepticons aided by the Third Reich.

During the Cybertronian civil war, the Autobots hired a yellow bot to lead them through the sea of rust to find the Allspark. However they soon learn that the yellow bot has secrets about himself, and why he knows his way around the sea so well.