- Saki Hyuga and Mai Mishou met for the first time when they were both nine years old, after they followed two glowing balls that flew towards the Sky Tree, a big tree situated atop a mountain in their town. At the same time, a boy named Hidari Tanichi met them on a night walk with a detective in a white coat. But they didn't expect that 5 years later, they would meet each other and that's when the story begins. A detective couple and a magical girl couple join forces to protect the Land of the Fountain and a city crying over mysterious cases.. - In Fuuto City, criminals using USBs called Gaia Memory are rampaging throughout the city with the cases the perpetrators have committed. This is when the two-in-one detective teams up with Pretty Cure to keep Fuuto smiling. - Come, settle your sins.