Bright and optimistic, Princess Poppy must save her fellow trolls from an Evil Bergen Chef, but not without the help of survivalist and the grumpiest troll, Branch.

When an evil spirit known as Pitch Black lays down the gauntlet to take over the world, immortal Guardians must join forces for the first time to protect the hopes, beliefs and imagination of children all over the world.

The gang are back and are still trying to make it back to New York's Central Park Zoo. They are forced to take a detour to Europe. When French animal-control officer Capitaine Chantel DuBois picks up their scent, Alice and company are forced to hide out in a traveling circus.

Down-on-his-luck private eye Eddie Valiant gets hired by cartoon producer R.K. Maroon to investigate an adultery scandal involving Jesse Rabbit, the handsome husband of Maroon's biggest star, Rosie Rabbit. But when Minerva Acme, Jesse's alleged side piece and the owner of Toontown, is found dead, the villainous Judge Doom vows to catch and destroy Rosie.

Construction Worker Emily Brickowski is mistakenly identified as the Special-- an extraordinary being and the key to saving the world.