When Decepticons escape to the world of Zootopia, Optimus Prime must follow them and prevent a new plot from both the locals and the Decepticons. Alone and caught in the middle of a hunt for his life-spark, he befriends a lonely jaguar orphan named Jake, who inadvertently holds the key to restoring the desolate Cybertron. Together they face the odds of Megatron's Decepticons and the challenge of keeping Prime a 'Robot in Disguise' from a whole world.