Transported from her summer home without a second warning, Oriana found herself in the midst of Red Grave City just as the Qliphoth Tree was maturing. Easily accepting her new situation, she brings up her sword and travels with the Devil May Cry as they scale the tree and attempt to save the city; as well as the world. Sticking out like a sore thumb is V, a mysterious man with demonic power that are unlike the others. To the outsider, he looks like a frail person. But to her, he was a reflection of her older self and someone with more depth than what one can see from his outward appearance. In the course of the journey, her curiosity has led her to be by his side, and to stay there, through thick and thin and through the shadows under veil of night.