Echo and Callie grew up with their many siblings, living in an open stretch of land behind Twolegplace. But one day, a forest fire happens. Cats living in the forest go to their home for refuge, with no knowledge of their family living there. They flee, but Echo and Callie get lost and separated from their family in the burning forest. They narrowly escape, but their lives take an unexpected turn neither of them would have ever thought before... While Echo works hard to prove herself to the cats who helped her and her sister, Callie is stuck around camp, and she's tired of it. Little did she know that she should have been savouring every moment...